1947a). A second collection of 13 fish in 1867 by Dr. Francis
Sumichrast, an ornithologist, came from Cosamaloapan on
the Rio Papaloapan, Veracruz, Mexico, where the Smithso-
nian had maintained a field station for several decades. The
third collection of 68 fish was made by Seth E. Meek in 1902
(Meek, 1904), near the railroad station of El Hule, Oaxaca,
Mexico, the last stop of the Transisthmian Railroad before
it passed over the Rio Papaloapan into the state of Veracruz.
Meek, an ichthyologist, had traveled by railroad from Chia-
pas in the south to the state of Nuevo Leon in the north. He
detrained every other stop to collect fishes and then con-
tinued his journey the next day. Myron, in his search for
platyfish, made the most sensible decision: he headed for
the Rio Papaloapan.
Gordons 1930 Mexico Expedition was financially sup-
ported by the National Research Council and by the Mu-
seum of Zoology of the University of Michigan, which
wanted to enlarge its fish collection. The expedition con-
sisted of three men: Gordon, Edward Creaser, a malacolo-
gist from Cornell, and Ricardo Ostos, M.D. They drove to
Laredo and had to ford the Rio Grande, then proceeded
through Monterrey along the foot of the Sierra Madre to
Ciudad Victoria. The section from Monterrey to Ciudad
Victoria was unpaved and every stream had to be forded.
They learned to wrap their socks around the sparkplugs to
keep them dry. This entire distance of 267 km took 10 hours
to drive. From Ciudad Victoria they continued to Ciudad
Valles and then headed southeast to Tantoyuca, crossing the
Rio Moctezuma at El Higo. Here they stayed for several days
on Ricardo Ostos ranch and explored the surrounding area
on horseback (Figure 1). One day, 20 km south of Tan-
toyuca, they collected 4 immature swordtails near the Rio
de los Hules and Rio Calaboza, which Gordon identified as
X. montezumae. Actually, these fish represent the first speci-
mens of X. birchmanni, a species not described until 1987.
No further collections of Xiphophorus were made in this
region until 1985. From Tantoyuca the expedition headed
east to the Estero Cucharas, a small stream flowing into the
Laguna Tamiahua, about 75 km south of Tampico. The
road conditions were so horrendous that they had to rest
for half an hour every 3 km. From there they turned south
toward the Rio Cazones and then climbed up over terrible
roads across the headwaters of the Rio Tecoluta to the
Mexican Plateau. The Rio Tecoluta watershed is character-
ized by steep canyons and waterfalls several hundred meters
high. In small streams on a narrow plateau above the falls at
1200-m altitude Dr. Gordon discovered a platyfish, some-
what similar to X. variatus, many years later described as X.
evelynae. A series of hydroelectric power plants and dams
had been constructed in this rather inaccessible region, and
undoubtedly Gordon was aided by Claudio Martinez, an
alumnus of Cornell and an engineer with the Mexican
Power Company. The expedition headed for Mexico City,
where they rested, and then drove to Veracruz.
On the way they collected swordtails, X. helleri, at
Jalapa and were puzzled by the many large males with in-
completely formed gonopodia and undifferentiated gonads
(Gordon, 1937a). The only record that I can find about
their next move states that they hired a boat at Alvarado and
headed up the Rio Papaloapan toward Cosamaloapan and
El Hule, 45 and 85 km, respectively, to the south, but more
than twice that distance by river. A railroad spur from Ve-
racruz ends at Alvarado, but I do not think they took the
train when they could have traveled on another one to El
Hule. It is also unlikely that they took the Ford there, be-
cause the track to Alvarado on the narrow high ground
between the high dunes to the left and the marshes to the
right must have been blocked by the ever-present sand
drifts. Gordon placed Alvarado only 20 miles from Vera-
cruz; however, it is actually twice that far. I think he may
have hired the launch at Boca del Rio, 20 km south of
Figure 1. Dr. Myron Gordon at Tantoyuca, Veracruz, Mexico,
1930.
The Xiphophorus Problem
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