Today, Tula is not considered such a bad place for
Xiphophorus researchers. Only 15 years ago we learned of a
level path through the barrier range that leads to a valley at
1000-m elevation, only 20-km distant, with a small stream
teaming with swordtails, X. nezahualcoyotl.
Early in April the expedition arrived in Veracruz, where
they stored the model T. From there they proceeded in
relative comfort by local train to El Hule, which had been
renamed Papaloapan. For a belated impression of the joys
of riding this train, the reader is referred to Theroux (1979).
Papaloapan lies near the fall line. To the east of the railroad
track stretches the low coastal plain with its enormous
swamps, muddy lagoons, and oxbows, and to the west we
find the beginning of the Piedmont of the Sierra Madre del
Sur. Today, a narrow vehicle bridge crosses the river at
Papaloapan with a toll station at its southern (Oaxaca) side.
Truck traffic on the road is as busy as on the New Jersey
Turnpike because it provides the least mountainous access
to Puebla and Mexico City from the southern states. A new
superhighway built through the marshes 30 km to the east
is barely used because of its excessive toll rates.
But in 1932 the area was a sleepy backwater. The Stan-
dard Fruit Company had banana plantations in this area
with wonderful facilities for its administrative staff. With
letters of introduction to military and civil authorities, the
second Cornell expedition was taken in by the Fruit Com-
pany and installed like royalty. After camping out for nearly
a month, it felt wonderful. The perfect service and comfort
of our headquarters meant more to us than a paid-up, de
luxe suite of rooms at Sun Valley Lodge (Gordon, 1940).
The companys officers must have looked on the three New
Yorkers as strange little fish, indeed, to have traveled 3000
miles to catch 1-inch-long fish! Every facility of the com-
pany was placed at Dr. Gordons disposal: vehicles, boats,
horses, and guides. Three days fishing effort yielded not a
single platyfish. Gordon had not yet discovered its exact
ecological niche. He thought platyfish, being rather broad
and somewhat deep-bodied, could not be a stream fish, but
must prefer quiet waters without current. So he concen-
trated on the nearby oxbows and swamps. From my own
experience, I know how utterly depressing it can be to have
come so far and fished for many days without catching a
single specimen. Myron must have felt likewise, but he
merely wrote they were quite discouraged. Eventually,
they headed upstream by boat from company headquarters
and entered the Rio Tonto and one of its little side streams.
And there Gordon caught his first X. maculatus!
Excitedly, and no doubt with satisfaction, he tele-
graphed his success to Cornell. On April 17, 1932, the New
York Times carried a long article with the headlines: Can-
cerous Fish Found in Mexico, Oaxaca Jungle Pierced. A
direct quotation from the telegram reads, We found by
actual count ninety-nine platyfish, some of which are sus-
piciously suffering with a small degree of melanosis. There
is one in which the condition is well seen.
And of course the publicity hounds of Cornell sprang
into action and added that the Cornell expedition attained
its objective, deep in the jungle recesses of the Mexican
state of Oaxaca. In reality, the jungle had been long gone
and the only Xiphophorus person, if we may call him that, to
rhapsodize about the luxuriant vegetation was C.B. Heller,
a 21-year-old German botanist who landed in Veracruz in
1845 (Siemens, 1990). One of his objectives had been to
scout for areas suitable for German colonists, and while
doing so he collected in the Rio Blanco drainage the first
swordtail, which now carries his name. This river is the
northernmost tributary to the Rio Papaloapan estuary, and
the general area where Heller found the swordtail is ap-
proximately 100 km northwest of the village where Gordon
caught his first platyfish.
The next day, the platyfish and also some swordtails
were placed in the cans and successfully shipped by railroad
to a commercial fish breeder in New Orleans. Later in the
summer when it was warm up north, they were reshipped to
Cornell. Melanotic platyfish, however, were not to be.
Closer examination later showed that the darker fish merely
had a well-expressed spot-sided pattern, and the single most
heavily pigmented one exhibited a new pattern, called spot-
ted-belly, Sb (Gordon and Smith, 1938a), or black-
bottomed (Gordon, 1948), but there was no melanosis. It
remains the only Sb fish that has ever been caught in the Rio
Papaloapan drainage, although fish with somewhat similar
patterns have been taken in some of the river systems to the
east (Kallman, 1975). Next day the expedition members
packed their bags and retraced their steps to Mexico City to
collect goodeid fishes on the Mexican Plateau to satisfy their
sponsors in the aquarium trade. Later, before the return trip
north, Myron inquired at the Mexican Automobile Asso-
ciation about the new road being constructed beyond the
old towns of Zimapan and Jacala to Tamazunchale at the
foot of the Sierra Madre; they were not going to pass
through Tula again. Assured that on average 12 cars per day
traveled over this road, the expedition opted for the new
highway (Gordon, 1933).
For 160 km beginning just beyond Zimapan the road
passes from 2000-m elevation through lush forests of pine,
The Xiphophorus Problem
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