chemicals was developed. This so-called medaka specific-
locus test system has led to more mechanistic studies. Suc-
cess in detecting AP-PCR/RAPD polymorphisms between
two inbred strains, one established from the northern popu-
lation of the Japanese wild medaka and the other from the
southern population, allowed these researchers to generate
a draft linkage map. This preliminary map will be followed
by a more definitive one (see Shima and Shimada, this
issue).
In 1999, the Environmental Agency of Japan listed the
Japanese medaka as an endangered species. Wild popula-
tions of medaka in Japan have been reduced by loss of
habitat, including irrigation canals, swamps, marshes, and
ponds. At present, conservation of the species tends to be
ignored. Shima and his colleagues are convinced that the
development of the medaka as a new model animal should
parallel the scientifically sound preservation and mainte-
nance of the species and its strains. At least 10 specimens
each from nearly all wild medaka populations in Japan are
being archived and preserved in 100% ethanol as DNA
specimens.
Services
The current stock resources of Japanese medaka in-
clude the following ones: Fourteen northern Japanese popu-
lations; 54 southern Japanese populations; 5 hybrid popu-
lations; 6 Chinese and Korean populations; 2 tester strains;
22 induced specific-locus mutant strains; and 2 congenic
strains. If the requested numbers of embryos or fish are
available, they are distributed to researchers as long as the
stocks are clearly used only for academic purposes and
proper credit for their origin is given in publications.
Contacts
Dr. Akihiro Shima: shima@biol.s.u-tokyo.ac.jp; De-
partment of Integrated Biosciences, Graduate School of
Frontier Sciences, The University of Tokyo, Tokyo, Japan
113-0033. Dr. Atsuko Shimada, Department of Biological
Sciences, School of Science, The University of Tokyo, To-
kyo, Japan 113-0033. Dr. Kenjiro Ozato, Yuko Wakamatsu,
Laboratory of Freshwater Fish Stocks, Nagoya University
Bioscience Center, Nagoya, Japan 464-8601; telephone and
fax +81 3 5802 2911.
AQUATIC RESOURCES AT THE UNIVERSITY
OF GEORGIA
Program Description
Transgenic Japanese medaka (Oryzias latipes) and mum-
michog (Fundulus heteroclitus) are new research resources
available at the Warnell School of Forest Resources, Uni-
versity of Georgia (Winn et al., 2000, this issue). These
transgenic fish were developed to improve methods used to
assess the health risks of exposure to chemicals in aquatic
environments and to provide new animal models to study
comparative in vivo mutagenesis. The transgenic fish carry
prokaryotic vectors, bacteriophage lambda LIZ, or plasmid
pUR288, harboring the mutational target genes (e.g.,
lambda lacI, lambda cII or LacZ) that are identical to those
employed in transgenic rodents. Using similar mutation
assay procedures, the target genes are separated from the
fish genomic DNA and transferred into specialized bacterial
host cells to quantify spontaneous and induced mutations.
Many fundamental parameters of mutagenesis are
similar in transgenic rodents and fish. Spontaneous mutant
frequencies are low in the fish models, comparable to the
ranges in transgenic rodents, which facilitates sensitive de-
tection of mutations. Exposing fish to different chemical
mutagens has demonstrated that their mutational targets
respond in a manner consistent with known mechanisms of
mutagen action resulting in concentration-dependent, tis-
sue-specific, and time-dependent mutations. Spontaneous
and induced mutational spectra in the fish also are consis-
tent with those of transgenic rodents. Therefore, in vivo
mutation analyses using transgenic fish are feasible and il-
lustrate the potential value of fish as comparative animal
models. The transgenic loci are genetically neutral, a feature
that is advantageous compared to other assays such as those
involving endogenous genes that may be limited to specific
tissues or developmental stages. As a result, mutations in
neutral transgenes accumulate and persist, avoiding the in-
fluence of selective pressures in vivo on mutational fre-
quency. Recent studies in rodents indicate that some, but
not all potent genotoxins can be detected after a single acute
treatment (Hoorn et al., 1993; Morrison and Ashby, 1994;
Dolle et al., 1996). Consequently, the more treatments that
are given, the more mutations will be induced, and the
more sensitive the test. The limitation here is that multiple
doses can be more toxic than single doses, often requiring a
compromise between number of treatments and the dosage.
S242
William E. Hawkins et al.
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